SUILLUS GLANDULOSIPES Thiers and Smith, Contr. Toward a Monogr. of N. Am. Sp. of Suillus, p. 86. 1964

Illustrations: See Microfiche No. 42

Pileus 6-12 cm broad at maturity, convex when young, becoming broadly convex to plano-convex to plane or shallowly depressed with age, often highly uneven to undulating or irregular toward the margin when old; surface viscid, when young glabrous or with irregularly distributed fibrils near the margin and more rarely on the disc, typically becoming glabrous with age, with the gluten drying in streaks, rarely appearing somewhat fibrillose and pitted or rugose when older; color at first yellow to pale tan or reddish cinnamon ("pale ochraceous-buff" to "pinkish buff" to "vinaceous-cinnamon" to "orange-cinnamon" to "cinnamon" to "apricot buff" to "ochraceous-orange"), unchanging with age or becoming reddish brown to brown ("ochraceous-tawny" to "Mikado brown" to "Verona brown" to "warm sepia" to "cinnamon-rufous"); margin strongly incurved to inrolled, with a conspicuous white cottony roll when young, disappearing with age, entire. Context 1-2 cm thick, white when very young, changing to pale yellow (near "marguerite yellow" to "cartridge buff" to "ivory yellow"), unchanging when bruised or rarely becoming pale lavender ("avellaneous") with age, firm. Taste and odor not distinctive.

Tubes 0.4-1 cm long, decurrent to subdecurrent to occasionally shallowly depressed, when young tan ("warm buff" to "light ochraceous-buff" to "pale ochraceous-buff" to "cream buff"), changing to yellow ("antimony yellow" to "chamois") with age, often with reddish brown glandulae on the pores, unchanging when bruised; pores angular, one to two per millimeter, concolorous, unchanging when bruised.

Stipe 4-11 cm long, 1-2 cm thick at the apex, with white mycelium at the base, equal, solid; surface dry, white to pale yellow ("marguerite yellow" to "primrose yellow") to occasionally pale pink ("pale pinkish buff"), unchanging with age, densely covered with irregularly shaped glandulae that are dark brown to black and often stain fingers dark brown when handled; no annulus. Context white when young, becoming tan to pale yellow ("warm buff" to "ivory yellow") with age, usually unchanging when exposed, but sometimes darkening slightly at the base, or becoming vinaceous in older basidiocarps.

Spore print brown. Spores 6-9(12) X 3-4 Ám, hyaline to pale ochraceous in KOH, smooth, thin-walled, cylindric to subellipsoid.

Basidia 23-28 X 5-8 Ám, clavate, four-spored, hyaline, contents often granulose in KOH. Hymenial cystidia 30-50 X 7-12 Ám, abundant only on the pores, fasciculate, incrustations on basal portion staining dark brown when revived in KOH, contents sometimes staining brownish, no solitary cystidia seen.

Tube trama hyaline, divergent from a distinct mediostratum, subgelatinous, hyphae up to 7 Ám wide, tissue at junction of pileus and tube trama often staining pale vinaceous in KOH. Pileus trama interwoven, homogeneous. Pileus cuticle differentiated as a broad (330 Ám) ixotrichodermium of interwoven hyphae, contents of hyphae appear brownish-granulose in KOH, outermost hyphae often somewhat enlarged and incrusted, hypodermium well differentiated as a compactly interwoven layer staining dark brown in KOH. Stipe cuticle differentiated as a gelatinous layer with numerous fascicles of darkly staining caulocystidia similar to the hymenial cystidia. Clamp connections absent.

Chemical reactions unknown.

Habit, habitat, and distribution Gregarious to cespitose in humus under Bishop and, less commonly, beach pine. In California, Suillus glandulosipes is known only from the northern coastal regions. It is relatively rare in California and, with the exception of one collection made in Patricks Point State Park in Humboldt County, it has been found only in Jackson State Forest in Mendocino County. Although originally described from California, it is now known from throughout the Pacific Northwest and as far east as Michigan.

Material studied Humboldt County: Thiers 14516. Mendocino County: Thiers 8394, 9307, 9332, 9335, 9472, 10702, 14150, 18106, 21414, 21443, 23063.

Observations Suillus glandulosipes is distinguished by the pale cinnamon pink to more vinaceous colored pileus, the large conspicuous glandulae on the stipe, and the well-developed, easily detected ring of velar tissue on the margin of the pileus. No evidence of any type of annulus has ever been observed. The ring of veil tissue is readily apparent on young basidiocarps but characteristically disappears with age. This species obviously belongs to the same group as S. granulatus and S. albidipes. The presence of the veil tissue in S. glandulosipes is perhaps the most significant difference, but the pale color of the pileus is also quite distinctive. Although it is indicated in the description that old basidiocarps may darken considerably, it should be emphasized that this is not the usual pattern. Suillus brevipes is commonly found in the same type of habitat and often fruits at the same time, but it characteristically has a much shorter stipe and lacks the noticeable glands on the surface.

Edibility not determined.

The Boletes of California
Copyright © 1975 by Dr. Harry D. Thiers
Additional content for the online edition © 1998 by Michael Wood, Fred Stevens, & Michael Boom
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