The North American Species of Pholiota

80. Pholiota ornatula (Murr.) comb. nov.

Gymnopilus ornatulus Murrill, Mycologia 4: 251. 1912.
Flammula ornatula Murrill, Mycologia 4: 262. 1912.

Illustrations: Text figs. 107-109.

Pileus caespitose, 3 cm broad, convex to nearly plane, gibbous or umbonate, slightly viscid when wet, fibrillose, flavo-melleous tinged with pale rose-brown, the latter color more conspicuous at the center.

Lamellae adnate, pallid when young, becoming pale-fulvous from the spores, plane, broad, of medium distance.

Stipe 5 cm long, 4 mm thick, smooth, glabrous and cremeous at the apex, subconcolorous and shaggy-fibrillose below.

Spores 5.5-7.5 x 3.5-4.5 µ, smooth, apical pore present and at times apex appearing slightly truncate, shape in face view elliptic to ovate, in profile obscurely inequilateral, pale tawny in KOH and only slightly paler in Melzer's reagent, walls 0.25 µ thick.

Basidia 18-22 x 5-6 µ, 4-spored, hyaline in KOH, yellowish in Melzer's reagent. Pleurocystidia present as imbedded chrysocystidia 18-25 x 6-10 µ, often 5-6 µ at basal septum, wall hyaline thin and smooth, content varying from a local area of granular-refractive material to the amorphous refractive mass characteristic of chrysocystidia and the inclusion merely yellowish in Melzer's reagent. Cheilocystidia 24-32 x 6-10 µ, subclavate to subcylindric or fusoid-ventricose.

Gill trama a central floccose strand of hyaline hyphae 3-9 µ broad with thin, smooth walls; subhymenium gelatinous, structure not distinct but apparently divergent-interwoven. Pileus cutis a gelatinous pellicle. No other data on type reliable.

Habit, Habitat, and Distribution: On soil, roadside, California, November. Type studied.

Observations: This species is near P. melliodora, in which the pileus margin remains incurved for a long time, but the cheilocystidia are longer and more slender, the subhymenium more distinctly gelatinous, and the gills are crowded and at first yellow. The odor in P. ornatula is not recorded.